京都大学 京都大学

Cell Wall Formation in Woody Plants

Numbering Code U-AGR05 3E135 LJ80 Year/Term 2022 ・ Second semester
Number of Credits 2 Course Type Lecture
Target Year 3rd year students Target Student
Language Japanese Day/Period Tue.2
Instructor name YOSHINAGA ARATA (Graduate School of Agriculture Associate Professor)
Outline and Purpose of the Course Focusing on the lignification within plant cell walls, we will discuss the mechanisms of differentiation into xylem cells and cell wall formation, the three-dimensional placement of cell walls at the molecular level, and the biosynthetic mechanism of cell wall components.
Course Goals Understand the basic structure, ultra-microstructure, chemical component distribution, and formation mechanism of cellulose - the important structural component of biomass.
Schedule and Contents Topics and class schedule are as follows: 1. Orientation (once)
　Overview of cell wall formation and an explanation of recent topics.

2. Cell segmentation and wood segmentation (once)
　The process of fragmentation in plant cells and the process by which undifferentiated cells differentiate into wood cells is explained.

3. Formation and structure of primary walls (once)
　The formation mechanism and structure of primary walls, and the three-dimensional placement and role of cell wall components at the molecular level is explained.

4. Formation and structure of secondary walls (twice)
The formation mechanism and structure of secondary walls, and the three-dimensional placement and role of cell walls at the molecular level is explained. In addition, we discuss the diversity of cell wall structures.

5. Biosynthetic, transported, and deposited cell wall components (six times)
　The biosynthesis, transport, and deposition processes of cellulose, hemicellulose, and lignin are explained. The mechanisms of deposition and assembly of these cell wall components are also described.

6. Modification of cell walls by GMOs (twice)
　The study examples of the modification of cell walls by genetic manipulation, which has been actively conducted in recent years, are introduced, and future possibilities are discussed.

7. Summary (once)
The series of lectures will be reviewed and supplementary explanations will be provided.

Regular examination

Feedback (once): During the feedback session, the instructor will be available in the laboratory to answer students' questions.
Evaluation Methods and Policy [Evaluation method]
Based on the results of the ordinary point evaluation (attendance and submission of tasks 20%) and the regular examination (written 80%).

[Evaluation criteria]
Evaluation standards and policies are based on the those described in the Faculty of Agriculture Handbook for the relevant year.
Course Requirements Nothing in particular
Study outside of Class (preparation and review) Separate instructions are given during class.
Textbooks Textbooks/References Not used. Lecture materials will be distributed.
References, etc. Nishitani, Toshiaki Umezawa, eds. "植物細胞壁" (Kodansha) ISBN: 978-4-06-153818-4, The Japan Wood Research Society ed., "木質の構造" (Buneido Publishing) ISBN: 978-4-8300-4120-4
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